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First encounter with Guardian Wraiths occurs at Vorago Solitude during the mission Beginning of the End. They also spawn in later discovered locations: Outfall Pumping Station, Tycho's Ribs and Eleseer.
Guardian Wraiths actively participate at boss battles with RK5 (with all other staff welding and flying Guardians).
Guardian Wraiths start their transformation when they first sight a target. They may alternate between two versions: Static Wraith (shock) and Corrosive Wraith, or Smoldering Wraith (fire) and Cryo Wraith.
- Guardian Wraiths (along with Guardian Cheru and Guardian Spectre) may be encountered before first enter in Lost Legion camp during story mission, at Eridian teleports not far from transition between Vorago Solitude and Triton Flats.
- Capturing of elemental charged Guardian Wraiths is an objective of optional mission Guardian Hunter.
- In True Vault Hunter Mode, when spawned, they are renamed Warden Wraith, and in Ultimate Vault Hunter Mode, they are renamed Fallen Wraith. Their elemental name prefixes change as follows: